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• This paper looks at the experience of gender mainstreaming in the Institutions for Inclusive Development (I4ID) programme – an adaptive, politically smart governance programme in Tanzania. • When development programmes try to engage with political stakeholders and align with the priorities of wider coalitions there is a danger that gender equality is de-prioritised. • It is important that formal political economy analysis, as well as other data collection, analysis and consultation exercises, are gender-sensitive. Teams should also look for ways to make gendered political and power analysis part of the everyday routine practice of staff. • Working politically and adaptively to advance gender objectives calls for staff with a specific skillset, as well as links to appropriate networks and political stakeholders. It also implies establishing checks and incentives to hold staff and partners accountable for gender objectives, and strong and consistent messaging from team leaders.